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Fragile Papershell

Fragile Papershell

North Dakota Species of Conservation Priority III

Fact Sheet
Scientific Name Leptodea fragilis
General Description Shell compressed and generally oblong. Both ends rounded with the exception of females where posterior end is expanded. Shell is commonly yellow to yellow/green with light green rays.
Status Year-round resident.
Abundance Common throughout its core range. Rare in North Dakota.
Primary Habitat Streams with mud, sand or gravel bottoms.
Federal Status None.
Reason for Designation Species was just recently document in the state. Little is known of the species and requires more investigation.


Fragile Papershell
Fragile Papershell

(Photos courtesy of Prairie Waters Education and Research Center - Valley City University)

Locations and Conditions of Key Habitat

Preferred Habitat

Species is found in varying stream sizes and bottom substrates throughout its range.

Key Areas and Conditions for Fragile Papershell in North Dakota

Documented only in the James River.

Problems Which May Affect this Species


Impoundment of the James River has changed the historic flow regime of the river. It has also blocked movements of fish species and in turn mussel species. Land use practices on surrounding lands have also impacted the river negatively. Chemical run-off and sedimentation contribute to deteriorated water quality.

Other Natural or Manmade Factors

No other threats have yet been identified for this species.

Research and Survey Efforts

Current Research or Surveys

  • No current research or survey efforts are on-going in this species range.

Previous Research or Surveys

  • Cvancara conducted a state-wide survey of the mussels of North Dakota in 1978.
  • The North Dakota Game and Fish Department revisited Cvancara’s sites in 1990.
  • Valley City State University revisited Cvancara’s sites and surveyed additional sites in 2008.
  • The NDDH conducted freshwater mussel surveys for state waters as a segment of its Index of Biotic Integrity (IBI) work.

Additional Research or Surveys Needed

  • Additional surveys of the James River should be conducted to find the range of this species.
  • A monitoring protocol for mussel species has been developed for the North Dakota Game and Fish Department under the SWG program. Implementation of this monitoring protocol is a future goal.

Management Recommendations

  • Develop buffers along riparian areas.
  • Work with partners to reduce the use of chemical near waterways.
  • Work with partners to reduce wetland drainage.
  • Remove river impoundments where possible.
  • Work with partners to maintain instream flows.

Monitoring Plans

A monitoring protocol has been developed for the North Dakota Game and Fish Department under the SWG program. Implementation of this monitoring protocol is a future goal.

2005-2015 Progress

Added as Level III Species of Conservation Priority in the 2015 revision of the Wildlife Action Plan. SWG T-24-R A Two Phase Population Survey of Mussels in North Dakota Rivers provided important information on the distribution of this species. Work to implement a monitoring protocol for mussels species will is a goal of the revised Wildlife Action Plan.

Range Map

Note: A listing of works consulted when compiling the information on this page may be found in the 2015 State Wildlife Action Plan.